Question: use of ~ in an input name.

In feature "Procedures for two animations" posted by Kitonium appears this line, within a procedure.

T1:=(O2*~y+O1*~u)/~(y+u): T2:=(O3*~u+O2*~v)/~(u+v): T3:=(O4*~v+O3*~z)/~(v+z):

What is the meaning of "~y", for instance, and how does it differ from "y"?

I understand the conventional usage of ~ with a name in output to imply that that name is subject to an assumption.

I have been able to find nothing pertinent in the Maple Help information.  Please enlighten me.

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